The "Regional Leadership in Wildland Fire Research Act of 2025" mandates the Secretary of Commerce, in coordination with the National Oceanic and Atmospheric Administration (NOAA) Administrator and the Chief of the Forest Service, to establish a competitive process for selecting and establishing regional wildland fire research centers. The Secretary must select at least seven institutions of higher education or land-grant universities, prioritizing those with existing wildland fire research programs, partnerships, or minority-serving institution status. These centers must be distributed across specific regions, including Alaska, California, the Northern Rockies, the Pacific Northwest, the Pacific Islands, the Southeast, and the Southwest. Each regional center will coordinate research with various entities to improve the understanding of wildland fire for management agencies. Their core functions include developing technologies and tools to predict fire potential, spread, behavior, and smoke impacts, as well as understanding vegetation response and post-fire risks. The centers are also tasked with developing career pathway training programs and implementing data management protocols that adhere to FAIR principles for open data exchange. A National Center Coordination Board will be established, composed of representatives from each regional center and co-chaired by the NOAA Administrator and the Forest Service Chief. This Board is responsible for setting wildland fire science and data management priorities, coordinating data collection, and preventing duplication of efforts among research entities. It will also support the adoption of new technologies and research findings by wildland fire management agencies. Furthermore, each regional center will have a Regional Advisory Board , comprising representatives from federal and state land management agencies, tribal organizations, and experts in wildland fire mitigation and management. These boards will ensure research coordination, communicate operational needs to the regional centers and the National Board, advise on research goals, and assist in disseminating research outputs and data to relevant agencies. The bill authorizes significant appropriations for the regional centers, starting at $60,000,000 for fiscal year 2026 and increasing annually through 2030, with an additional $1,000,000 per year for the National Center Coordination Board. These funds can be used for establishing and administering the centers, including construction and equipment acquisition. The Secretary is required to submit regular reports to Congress detailing the progress of the regional centers and offering recommendations for improving wildland fire research.